Meis2 Meis homeobox 2 [Mus musculus (house mouse)] - Gene

Summary

Official Symbol: Meis2provided by MGI
Official Full Name: Meis homeobox 2provided by MGI
Primary source: MGI:MGI:108564
See related: Ensembl:ENSMUSG00000027210 AllianceGenome:MGI:108564
Gene type: protein coding
RefSeq status: REVIEWED
Organism: Mus musculus
Lineage: Eukaryota; Metazoa; Chordata; Craniata; Vertebrata; Euteleostomi; Mammalia; Eutheria; Euarchontoglires; Glires; Rodentia; Myomorpha; Muroidea; Muridae; Murinae; Mus; Mus
Also known as: Mrg1; Stra10; A430109D20Rik
Summary: This gene encodes a homeobox protein belonging to the TALE ('three amino acid loop extension') family of homeodomain-containing proteins. TALE homeobox proteins are highly conserved transcriptional regulators and several members have been shown to be essential contributors to developmental programs. In mice, a knock-out of this gene leads to lethality at embryonic day 14, accompanied with hemorrhaging. Embryos lacking this gene show defects in tissues derived from the neural crest, suggesting a critical role of this gene during cranial and cardiac neural crest cell development. Alternative splicing results in multiple transcript variants. [provided by RefSeq, Sep 2016]
Expression: Broad expression in frontal lobe adult (RPKM 31.3), CNS E14 (RPKM 21.9) and 17 other tissues See more
Orthologs: human all
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Genomic context

Location:: 2 E4- E5; 2 58.28 cM

Exon count:: 19

Annotation release	Status	Assembly	Chr	Location
RS_2024_02	current	GRCm39 (GCF_000001635.27)	2	NC_000068.8 (115691745..115897925, complement)
108.20200622	previous assembly	GRCm38.p6 (GCF_000001635.26)	2	NC_000068.7 (115861264..116067537, complement)

Chromosome 2 - NC_000068.8 Genomic Context describing neighboring genes

Genomic regions, transcripts, and products

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Go to reference sequence details

Genomic Sequence:

Go to nucleotide: Graphics FASTA GenBank

Expression

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See details

Project title: Mouse ENCODE transcriptome data
Description: RNA profiling data sets generated by the Mouse ENCODE project.
BioProject: PRJNA66167
Publication: PMID 25409824
Analysis date: n/a

Bibliography

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GeneRIFs: Gene References Into Functions

What's a GeneRIF?

Touch receptor end-organ innervation and function require sensory neuron expression of the transcription factor Meis2.
Title: Touch receptor end-organ innervation and function require sensory neuron expression of the transcription factor Meis2.
The neuronal transcription factor MEIS2 is a calpain-2 protease target.
Title: The neuronal transcription factor MEIS2 is a calpain-2 protease target.
The autism-associated Meis2 gene is necessary for cardiac baroreflex regulation in mice.
Title: The autism-associated Meis2 gene is necessary for cardiac baroreflex regulation in mice.
Dlx1/2-dependent expression of Meis2 promotes neuronal fate determination in the mammalian striatum.
Title: Dlx1/2-dependent expression of Meis2 promotes neuronal fate determination in the mammalian striatum.
Neural crest cells require Meis2 for patterning the mandibular arch via the Sonic hedgehog pathway.
Title: Neural crest cells require Meis2 for patterning the mandibular arch via the Sonic hedgehog pathway.
Control of mouse limb initiation and antero-posterior patterning by Meis transcription factors.
Title: Control of mouse limb initiation and antero-posterior patterning by Meis transcription factors.
Axial skeleton anterior-posterior patterning is regulated through feedback regulation between Meis transcription factors and retinoic acid.
Title: Axial skeleton anterior-posterior patterning is regulated through feedback regulation between Meis transcription factors and retinoic acid.
The transcriptional regulator MEIS2 sets up the ground state for palatal osteogenesis in mice.
Title: The transcriptional regulator MEIS2 sets up the ground state for palatal osteogenesis in mice.
Methylation impairs interaction with CRM1 without affecting PBX1 dimerization and thereby allows MEIS2 nuclear accumulation, a prerequisite for neuronal differentiation.
Title: Arginine Methylation Regulates MEIS2 Nuclear Localization to Promote Neuronal Differentiation of Adult SVZ Progenitors.
a previously unknown link between PcG proteins and downregulation of RA-related signals to mediate the phase transition of Meis2 transcriptional status during forelimb patterning, is reported.
Title: Variant PRC1 competes with retinoic acid-related signals to repress Meis2 in the mouse distal forelimb bud.

Submit: New GeneRIF Correction See all GeneRIFs (26)

Variation

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Alleles

Alleles of this type are documented at Mouse Genome Informatics (MGI)

Targeted (7) 1 citation
Endonuclease-mediated (2) 1 citation

Interactions

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Products	Interactant	Other Gene	Complex	Source	Pubs	Description

General gene information

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Markers

RH136240 (e-PCR)
- UniSTS:210234

AU048515 (e-PCR)
- UniSTS:144965

RH138147 (e-PCR)
- UniSTS:220234

Meis2 (e-PCR), detects polymorphism
- UniSTS:531343

Mrg1 (e-PCR), detects polymorphism
- UniSTS:507072

D2Mit132 (e-PCR), detects polymorphism
- UniSTS:128992

RH132496 (e-PCR)
- UniSTS:215779

WI-18493 (e-PCR)
- UniSTS:50670

UniSTS:234491 (e-PCR)
- UniSTS:234491

NoName (e-PCR)
- UniSTS:234492

Gene Ontology Provided by MGI

Function	Evidence Code	Pubs
enables DNA binding	IDA Inferred from Direct Assay more info	PubMed
enables DNA binding	ISO Inferred from Sequence Orthology more info
enables DNA-binding transcription activator activity, RNA polymerase II-specific	IBA Inferred from Biological aspect of Ancestor more info
enables DNA-binding transcription activator activity, RNA polymerase II-specific	IDA Inferred from Direct Assay more info	PubMed
enables DNA-binding transcription activator activity, RNA polymerase II-specific	ISO Inferred from Sequence Orthology more info
enables RNA polymerase II cis-regulatory region sequence-specific DNA binding	IBA Inferred from Biological aspect of Ancestor more info
enables RNA polymerase II cis-regulatory region sequence-specific DNA binding	ISO Inferred from Sequence Orthology more info
enables sequence-specific DNA binding	IDA Inferred from Direct Assay more info	PubMed
enables sequence-specific DNA binding	IGI Inferred from Genetic Interaction more info	PubMed
enables sequence-specific DNA binding	ISO Inferred from Sequence Orthology more info
enables sequence-specific double-stranded DNA binding	ISO Inferred from Sequence Orthology more info
enables transcription factor binding	IDA Inferred from Direct Assay more info	PubMed

Process	Evidence Code	Pubs
involved_in animal organ morphogenesis	IBA Inferred from Biological aspect of Ancestor more info
involved_in brain development	IBA Inferred from Biological aspect of Ancestor more info
involved_in embryonic pattern specification	IBA Inferred from Biological aspect of Ancestor more info
involved_in eye development	IBA Inferred from Biological aspect of Ancestor more info
acts_upstream_of_or_within eye development	IGI Inferred from Genetic Interaction more info	PubMed
acts_upstream_of_or_within multicellular organism development	IEA Inferred from Electronic Annotation more info
involved_in negative regulation of myeloid cell differentiation	IDA Inferred from Direct Assay more info	PubMed
involved_in positive regulation of cardiac muscle myoblast proliferation	ISO Inferred from Sequence Orthology more info
involved_in positive regulation of cell population proliferation	IBA Inferred from Biological aspect of Ancestor more info
involved_in positive regulation of mitotic cell cycle	ISO Inferred from Sequence Orthology more info
involved_in positive regulation of transcription by RNA polymerase II	IBA Inferred from Biological aspect of Ancestor more info
involved_in positive regulation of transcription by RNA polymerase II	ISO Inferred from Sequence Orthology more info
involved_in regulation of DNA-templated transcription	IEA Inferred from Electronic Annotation more info
acts_upstream_of_or_within visual learning	IMP Inferred from Mutant Phenotype more info	PubMed

Component	Evidence Code	Pubs
located_in cytoplasm	IEA Inferred from Electronic Annotation more info
is_active_in nucleus	IBA Inferred from Biological aspect of Ancestor more info
located_in nucleus	IDA Inferred from Direct Assay more info	PubMed

General protein information

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Preferred Names: homeobox protein Meis2

Names: Cbp/p300-interacting transactivator with Glu/Asp-rich carboxy-terminal domain 2; meis1-related protein 1; myeloid ecotropic viral integration site-related gene 1

NCBI Reference Sequences (RefSeq)

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RefSeqs maintained independently of Annotated Genomes

These reference sequences exist independently of genome builds. Explain

These reference sequences are curated independently of the genome annotation cycle, so their versions may not match the RefSeq versions in the current genome build. Identify version mismatches by comparing the version of the RefSeq in this section to the one reported in Genomic regions, transcripts, and products above.

mRNA and Protein(s)

NM_001136072.2 → NP_001129544.1 homeobox protein Meis2 isoform 1

See identical proteins and their annotated locations for NP_001129544.1

Status: REVIEWED

Description

Transcript Variant: This variant (1) differs in the 5' and 3' UTRs and has multiple differences in the coding region, compared to variant 9. The encoded isoform (1) is shorter and has distinct N- and C-termini compared to isoform 9.

Source sequence(s)

AK146633, AL929037

Consensus CDS

CCDS50666.1

UniProtKB/TrEMBL

Q3TY73, Q3UJ35

Related

ENSMUSP00000106533.3, ENSMUST00000110908.9

Conserved Domains (2) summary

pfam05920
Location:294 → 333

Homeobox_KN; Homeobox KN domain

pfam16493
Location:110 → 194

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
NM_001159567.1 → NP_001153039.1 homeobox protein Meis2 isoform 3

See identical proteins and their annotated locations for NP_001153039.1

Status: REVIEWED

Description

Transcript Variant: This variant (3) differs in the 5' and 3' UTRs and has multiple differences in the coding region, compared to variant 9. The encoded isoform (3) is shorter and has distinct N- and C-termini compared to isoform 9.

Source sequence(s)

AK146633, AK158839, AL929037

Consensus CDS

CCDS50668.1

UniProtKB/TrEMBL

Q3TY73

Related

ENSMUSP00000106532.2, ENSMUST00000110907.8

Conserved Domains (2) summary

pfam05920
Location:294 → 333

Homeobox_KN; Homeobox KN domain

pfam16493
Location:110 → 194

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
NM_001159568.1 → NP_001153040.1 homeobox protein Meis2 isoform 4

See identical proteins and their annotated locations for NP_001153040.1

Status: REVIEWED

Description

Transcript Variant: This variant (4) differs in the 5' UTR, and has multiple differences in the coding region, compared to variant 9. The encoded isoform (4) is shorter and has a distinct N-terminus compared to isoform 9.

Source sequence(s)

AK146633, AL929037, CX568764

Consensus CDS

CCDS50667.1

UniProtKB/Swiss-Prot

O35676, O35677, P97367, P97403, P97404

UniProtKB/TrEMBL

Q3TY73

Related

ENSMUSP00000028639.7, ENSMUST00000028639.13

Conserved Domains (2) summary

pfam05920
Location:294 → 333

Homeobox_KN; Homeobox KN domain

pfam16493
Location:110 → 194

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
NM_001159569.2 → NP_001153041.1 homeobox protein Meis2 isoform 5

See identical proteins and their annotated locations for NP_001153041.1

Status: REVIEWED

Description

Transcript Variant: This variant (5) has multiple differences in the coding region and differs in the 3' UTR, compared to variant 9. The encoded isoform (5) is shorter and has a distinct C-terminus compared to isoform 9.

Source sequence(s)

AK146633, AK158839, AL929037, CJ143034

Consensus CDS

CCDS50665.1

UniProtKB/TrEMBL

B1AWK4, Q3TY73

Related

ENSMUSP00000106531.3, ENSMUST00000110906.9

Conserved Domains (2) summary

pfam05920
Location:293 → 332

Homeobox_KN; Homeobox KN domain

pfam16493
Location:109 → 193

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
NM_001159570.2 → NP_001153042.1 homeobox protein Meis2 isoform 6

See identical proteins and their annotated locations for NP_001153042.1

Status: REVIEWED

Description

Transcript Variant: This variant (6) has multiple differences in the coding region and differs in the 3' UTR, compared to variant 9. The encoded isoform (6) is shorter and has a distinct C-terminus compared to isoform 9.

Source sequence(s)

AK146633, AK158839, AL929037, CJ143034

Consensus CDS

CCDS50664.1

UniProtKB/TrEMBL

Q3TY73, Q3TYM2

Related

ENSMUSP00000073898.7, ENSMUST00000074285.8

Conserved Domains (2) summary

pfam05920
Location:293 → 332

Homeobox_KN; Homeobox KN domain

pfam16493
Location:109 → 193

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
NM_001346036.1 → NP_001332965.1 homeobox protein Meis2 isoform 9

Status: REVIEWED

Description

Transcript Variant: This variant (9) encodes the longest isoform (9).

Source sequence(s)

AK146633, AK158839, AL772135, AL929037, BB612458, BQ572308, CX568764

UniProtKB/TrEMBL

Q3TY73, Q3UJ35
NM_001346038.1 → NP_001332967.1 homeobox protein Meis2 isoform 10

Status: REVIEWED

Description

Transcript Variant: This variant (10) lacks an alternate exon in the coding region, compared to variant 9. The encoded isoform (10) is shorter than isoform 9.

Source sequence(s)

AK146633, AK158839, AL772135, AL929037, BB612458, CX568764

UniProtKB/TrEMBL

Q3TY73

Conserved Domains (2) summary

pfam05920
Location:293 → 332

Homeobox_KN; Homeobox KN domain

pfam16493
Location:109 → 193

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
NM_001346041.1 → NP_001332970.1 homeobox protein Meis2 isoform 11

Status: REVIEWED

Description

Transcript Variant: This variant (11) has multiple differences in the coding region, compared to variant 9. The encoded isoform (11) is shorter than isoform 9.

Source sequence(s)

AK146633, AK158839, AL772135, AL929037, BB612458, KM974644

UniProtKB/TrEMBL

Q3TY73

Conserved Domains (2) summary

pfam05920
Location:293 → 332

Homeobox_KN; Homeobox KN domain

pfam16493
Location:109 → 193

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
NM_001346044.1 → NP_001332973.1 homeobox protein Meis2 isoform 12

Status: REVIEWED

Description

Transcript Variant: This variant (12) has multiple differences in the coding region and differs in the 3' UTR, compared to variant 9. The encoded isoform (12) is shorter and has a distinct C-terminus, compared to isoform 9.

Source sequence(s)

AK146633, AK158839, AL772135, AL929037, BB612458, BQ572308

UniProtKB/TrEMBL

Q3TY73
NM_001346045.1 → NP_001332974.1 homeobox protein Meis2 isoform 7

Status: REVIEWED

Description

Transcript Variant: This variant (7) differs in the 5' and 3' UTRs and has multiple differences in the coding region, compared to variant 9. The encoded isoform (7) is shorter and has distinct N- and C-termini compared to isoform 9.

Source sequence(s)

AK146633, AL772135, AL929037, BQ572308

UniProtKB/TrEMBL

Q3TY73
NM_001346047.1 → NP_001332976.1 homeobox protein Meis2 isoform 8

Status: REVIEWED

Description

Transcript Variant: This variant (8) differs in the 5' and 3' UTRs and has multiple differences in the coding region, compared to variant 9. The encoded isoform (8) is shorter and has distinct N- and C-termini compared to isoform 9.

Source sequence(s)

AK012325, AK146633, AK158839, AL772135, AL929037, BY012436

UniProtKB/TrEMBL

Q3TY73

Conserved Domains (2) summary

pfam05920
Location:313 → 352

Homeobox_KN; Homeobox KN domain

pfam16493
Location:129 → 213

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
NM_001346053.1 → NP_001332982.1 homeobox protein Meis2 isoform 13

Status: REVIEWED

Description

Transcript Variant: This variant (14) differs in the 5' and 3' UTRs and has multiple differences in the coding region, compared to variant 9. The encoded isoform (13) is shorter and has distinct N- and C-termini compared to isoform 9. Variants 13 and 14 encode the same isoform (13).

Source sequence(s)

AK146633, AK158839, AL929037, BY290986

UniProtKB/TrEMBL

A0A0A7EPH8
NM_001346055.1 → NP_001332984.1 homeobox protein Meis2 isoform 13

Status: REVIEWED

Description

Transcript Variant: This variant (13) differs in the 5' UTR, uses a downstream start codon, has multiple differences in the coding region and differs in the 3' UTR, compared to variant 9. The encoded isoform (13) is shorter and has distinct N- and C-termini compared to isoform 9. Variants 13 and 14 encode the same isoform (13).

Source sequence(s)

AK146633, AK158839, AL929037, BY290986, CJ143034

UniProtKB/TrEMBL

A0A0A7EPH8
NM_001346056.1 → NP_001332985.1 homeobox protein Meis2 isoform 15

Status: REVIEWED

Description

Transcript Variant: This variant (15) differs in the 5' and 3' UTRs and has multiple differences in the coding region, compared to variant 9. The encoded isoform (15) is shorter and has distinct N- and C-termini compared to isoform 9.

Source sequence(s)

AK146633, AL929037, BY290986

UniProtKB/TrEMBL

A0A0A7EPH8

Conserved Domains (2) summary

pfam05920
Location:206 → 245

Homeobox_KN; Homeobox KN domain

pfam16493
Location:22 → 106

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
NM_010825.3 → NP_034955.1 homeobox protein Meis2 isoform 2

See identical proteins and their annotated locations for NP_034955.1

Status: REVIEWED

Description

Transcript Variant: This variant (2) differs in the 5' UTR, and has multiple differences in the coding region, compared to variant 9. The encoded isoform (2) is shorter and has a distinct N-terminus compared to isoform 9.

Source sequence(s)

AK146633, AL929037, KM974644

Consensus CDS

CCDS16568.1

UniProtKB/TrEMBL

Q3TY73, Q6GU28

Related

ENSMUSP00000099597.4, ENSMUST00000102538.11

Conserved Domains (2) summary

pfam05920
Location:294 → 333

Homeobox_KN; Homeobox KN domain

pfam16493
Location:110 → 194

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1

RefSeqs of Annotated Genomes: GCF_000001635.27-RS_2024_02

The following sections contain reference sequences that belong to a specific genome build. Explain

Reference GRCm39 C57BL/6J

Genomic

NC_000068.8 Reference GRCm39 C57BL/6J

Range

115691745..115897925 complement

Download

GenBank, FASTA, Sequence Viewer (Graphics)

mRNA and Protein(s)

XM_036159069.1 → XP_036014962.1 homeobox protein Meis2 isoform X12

Conserved Domains (2) summary

pfam05920
Location:148 → 187

Homeobox_KN; Homeobox KN domain

pfam16493
Location:1 → 48

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
XM_036159067.1 → XP_036014960.1 homeobox protein Meis2 isoform X11

Conserved Domains (2) summary

pfam05920
Location:148 → 187

Homeobox_KN; Homeobox KN domain

pfam16493
Location:1 → 48

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
XM_030248227.2 → XP_030104087.1 homeobox protein Meis2 isoform X13

Conserved Domains (2) summary

pfam05920
Location:148 → 187

Homeobox_KN; Homeobox KN domain

pfam16493
Location:1 → 48

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
XM_036159066.1 → XP_036014959.1 homeobox protein Meis2 isoform X10

Conserved Domains (2) summary

pfam05920
Location:211 → 250

Homeobox_KN; Homeobox KN domain

pfam16493
Location:1 → 48

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
XM_030248221.2 → XP_030104081.1 homeobox protein Meis2 isoform X5

Conserved Domains (2) summary

pfam05920
Location:313 → 352

Homeobox_KN; Homeobox KN domain

pfam16493
Location:129 → 213

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
XM_006498871.5 → XP_006498934.1 homeobox protein Meis2 isoform X9

See identical proteins and their annotated locations for XP_006498934.1

UniProtKB/TrEMBL

Q3TY73

Conserved Domains (2) summary

pfam05920
Location:313 → 352

Homeobox_KN; Homeobox KN domain

pfam16493
Location:129 → 213

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
XM_030248224.2 → XP_030104084.1 homeobox protein Meis2 isoform X9

UniProtKB/TrEMBL

Q3TY73

Conserved Domains (2) summary

pfam05920
Location:313 → 352

Homeobox_KN; Homeobox KN domain

pfam16493
Location:129 → 213

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
XM_036159058.1 → XP_036014951.1 homeobox protein Meis2 isoform X4

Conserved Domains (2) summary

pfam05920
Location:313 → 352

Homeobox_KN; Homeobox KN domain

pfam16493
Location:129 → 213

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
XM_036159055.1 → XP_036014948.1 homeobox protein Meis2 isoform X2

Conserved Domains (2) summary

pfam05920
Location:376 → 415

Homeobox_KN; Homeobox KN domain

pfam16493
Location:129 → 213

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
XM_036159064.1 → XP_036014957.1 homeobox protein Meis2 isoform X7

UniProtKB/TrEMBL

Q3TY73

Conserved Domains (2) summary

pfam05920
Location:376 → 415

Homeobox_KN; Homeobox KN domain

pfam16493
Location:129 → 213

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
XM_011239337.4 → XP_011237639.1 homeobox protein Meis2 isoform X1

See identical proteins and their annotated locations for XP_011237639.1

Conserved Domains (2) summary

pfam05920
Location:376 → 415

Homeobox_KN; Homeobox KN domain

pfam16493
Location:129 → 213

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
XM_036159061.1 → XP_036014954.1 homeobox protein Meis2 isoform X6

UniProtKB/TrEMBL

Q3TY73

Conserved Domains (2) summary

pfam05920
Location:376 → 415

Homeobox_KN; Homeobox KN domain

pfam16493
Location:129 → 213

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
XM_036159065.1 → XP_036014958.1 homeobox protein Meis2 isoform X8

UniProtKB/TrEMBL

Q3TY73

Conserved Domains (2) summary

pfam05920
Location:357 → 396

Homeobox_KN; Homeobox KN domain

pfam16493
Location:110 → 194

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
XM_006498864.5 → XP_006498927.1 homeobox protein Meis2 isoform X3

Conserved Domains (2) summary

pfam05920
Location:357 → 396

Homeobox_KN; Homeobox KN domain

pfam16493
Location:110 → 194

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
XM_030248225.2 → XP_030104085.1 homeobox protein Meis2 isoform X11

Conserved Domains (2) summary

pfam05920
Location:148 → 187

Homeobox_KN; Homeobox KN domain

pfam16493
Location:1 → 48

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1
XM_030248226.2 → XP_030104086.1 homeobox protein Meis2 isoform X12

Conserved Domains (2) summary

pfam05920
Location:148 → 187

Homeobox_KN; Homeobox KN domain

pfam16493
Location:1 → 48

Meis_PKNOX_N; N-terminal of Homeobox Meis and PKNOX1